Skeletal microstructure of helens, lateral spines of hyolithids
In hyolithids the skeleton consists of four elements: a conch, an operculum and a pair of long, logarithmically curved 'spines' called helens. These last elements are rarely preserved, and have therefore remained poorly known and enigmatic. We have studied three-dimensionally preserved helens of the species 'Hyolithes'lanceolatus, from the Permian of New South Wales, Australia, and 'Hyolithes'groenwalli, from the Cambrian of Bornholm, Denmark. Helens were massive calcareous elements.