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10.1111/j.1475-4983.2005.00445.x
ANTOÑANZAS, R., SEN, S. 2005. New species of Paraphiomys (Rodentia, Thryonomyidae) from the Lower Miocene of As-Sarrar, Saudia Arabia. Palaeontology, 48, 2, 223–233.
Raquel López Antoñanzas and Sevket Sen The family Thryonomyidae is represented in the Lower Miocene of Saudi Arabia by a single species, Paraphiomys knolli sp. nov. This new taxon differs from all other thryonomyids in being small, lower molars having a short metalophulid II and an isolated anterolabial cuspid, and upper molars being antero-posteriorly compressed and pentalophodont. A cladistic analysis involving all extinct and extant species of thryonomyids is provided. Paraphiomys knolli branches as the sister-species of Paraphiomys pigotti, type species of the genus. Wiley Online Library