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Article: Occurrences of the cool-water dalmanelloid brachiopod Heterorthina in the Upper Ordovician of North America

Papers in Palaeontology - Vol. 1 Part 3 - Cover Image
Publication: Papers in Palaeontology
Volume: 1
Part: 3
Publication Date: August 2015
Page(s): 237 253
Author(s): Jisuo Jin and David A. T. Harper
DOI: 10.1002/spp2.1014
Addition Information

How to Cite

JIN, J. and HARPER, D.A.T. 2015. Papers in Palaeontology. 1, 3, 237–253. doi: 10.1002/spp2.1014

Author Information

  • Jisuo Jin - Department of Earth Sciences, University of Western Ontario, London, Ontario, Canada (email:
  • David A. T. Harper - Palaeoecosystems Group, Department of Earth Sciences, Durham University, Durham, UK

Publication History

  • Issue published online: 10 AUG 2015
  • Article first published online: 5 MAY 2015
  • Manuscript Accepted: 16 MAR 2015
  • Manuscript Received: 9 FEB 2015

Funded By

Natural Science and Engineering Research Council of Canada
Danish Council for Independent Research

Online Version Hosted By

Wiley Online Library
Get Article: Wiley Online Library [Pay-to-View Access] |


Through a comparative study with the British type material, several dalmanelloid brachiopod species from the Upper Ordovician of the Cincinnati Arch area of North America (Laurentia) are now confirmed to belong to Heterorthina Bancroft. These include several species previously treated variously as Dalmanella or Onniella, including such classic and widely cited species as ‘Dalmanella emacerata (Hall) and Dalmanella fairmountensis Foerste. These North American forms of Heterorthina belong to the dalmanelloid clade that is characterized by a dorsal medial interspace, and of cool to cold-water origin. Outside of Laurentia, Heterorthina occurs predominantly in Avalonia and peri-Gondwanan terranes, such as Iberia (Spain), Armorica (France) and Bohemia, palaeogeographically located between 40° and 60° south. The episodic occurrences of several Heterorthina species in the lower to middle Katian of the Cincinnati Arch area, located around 20°S in the Late Ordovician, are interpreted as opportunistic invasions of a cool-water brachiopod fauna from higher palaeolatitudes, associated either with oceanic cooling events or with relatively cool deep-water mass incursions during marine transgressions.

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