A model for the morphogenesis of ribs in ammonites inferred from associated microsculptures
Eight morphological types of microsculpture have been recognized on the outer shell surface of well-preserved ammonites. They consist mainly of wrinkles and creases which were developed on a pliant material. They are interpreted as the result of compressive stresses which occurred on a free (i.e. uncalcified) periostracum. This is the first evidence of periostracal development in ammonoids, and its relation to the shell suggests a mode of calcification similar to present-day molluscs. This periostracum was also attached to the soft body along longitudinal lines.