Taxonomy, phylogeny, and variability of Pseudisograptus Beavis
Manubriate isograptids from the Castlemainian and Yapeenian (Arenig) of New Zealand, Australian, and Chinese collections are described and analysed in terms of population systematics. Pseudisograptus manubriatus (T. S. Hall, 1914) is divided into five subspecies, of which P. m. koi, P. m. harrisi, P. m. janus, and P. m. texanus are new; P. bellulus sp. nov. and five other Pseudisograptus species are described. A tentative phylogenetic scheme is constructed from their stratigraphic occurrence and a cladogram of pseudisograptids and other forms with manubriate features.