The external surface of Dictyonella and of other pitted brachiopods
The external surface of Dictyonella is characterized by a smooth depressed umbonal region and a pitted ornament. The exposed part of the smooth plate, here termed the colleplax, is enlarged by resorption of the outer shell. A model is developed to explain the mode of function of the colleplax and its relationships to the soft tissue. The apparently complicated surface pattern of Dictyonella and other pitted brachiopods is shown to be produced by simple radial growth modified by various factors.